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Most pattern mutations
in the Cornsnake are simple traits, inherited in typical Mendelian recessive
fashion. In at least two cases, the traits are involved with each other in
some way, although the mechanism is not fully understood. Combining pattern
mutations with color mutations can lead to some exceptional looking specimens
indeed. Since pattern mutations essentially affect the distribution of the
pigments, many specimens appear much brighter or cleaner than typically
patterned individuals of the same color morph.
Motley
This, the first recessive pattern trait discovered in cornsnakes, is also one of
the most complex. The mechanisms involved in the redistribution of
pattern are not understood at all. However, a few plausible theories
have been postulated. The motley pattern affects the entire
animal. Belly patterning is completely absent. In most examples
there is a complete dark line along the edge of the ventral scales, matching
in coloration what one would suppose the ventral checkering to be. The
supposition is that the ventral patterning has been aligned along the edges of
the ventral scales. Lateral pattern is either absent or has been reduced
to faint irregular striping along the lower sides. The dorsal pattern
may consist of either widely spaced elongated blotches, often fused at the
sides to form a ladder pattern, or nearly full-length parallel stripes. Extremely
variable, leading to several named variations such as
Q-Tipped,
Pin-Striped,
and
Hurricane.
It is known that the motley and stripe allele share the same
locus, yet the Motley appearance is dominant to that of Striped. Thus
specimens with two motley alleles appear motley patterned, those with two
striped alleles appear striped and those with one of each allele appear
Motley, but are heterozygous for Stripe.
Striped
Two nearly perfect
full-length dark stripes surround a full-length light mid-dorsal stripe.
Lateral pattern is largely reduced to fain irregular stripes or elongated
blotches. Often the pattern of stripes is broken near the vent and
onto the tail. As in the motley pattern variant, ventral pattern is
absent. Consistency of striping is extremely variable, with some
specimens appearing more blotched than striped. This has led to a variety of
named appearances - none of which seem to breed true in a consistent
fashion. Among these are
Cubed,
Patternless
and
Vanishing Stripe.
It is known that the motley and stripe allele share the same
locus, yet the Motley appearance is dominant to that of Striped. Thus
specimens with two motley alleles appear motley patterned, those with two
striped alleles appear striped and those with one of each allele appear
Motley, but are heterozygous for Stripe.
Zigzag or Zipper
Appears to be caused by a longitudinal misalignment of the two
(left and right) halves of the dorsal pattern blotches. The result
erratic combinations often form a broken pattern resembling a thick wavy line
down the dorsum. A Zigzag or Zipper, if you will. Although the
trait is a simple recessive, specimens which are known homozygous for the
trait occasionally fail to exhibit it visually. In actuality, it just
happens that the misalignment of the two may be re-aligned by chance - and the
result appears completely normal to our eyes. Most examples of this
mutation have areas which appear completely normal due to this.
Recently, breeders have concentrated on this trait and have 'enhanced' it into
more and more severe pattern deformations. As they are stabilized genetically,
many are given new trade names. Aztec is one we hear a lot.
Plain-Belly
A total lack of ventral pattern is found in another cornsnake
color mutation. The early examples of the 'Blood-Red' morph lacked all
traces of ventral patterns. It has been demonstrated that this trait is
inherited separately from the blood-red trait, and can be passed to other
color mutations. It has also been conclusively demonstrated that it is genetically
distinct from the lack of ventral pattern exhibited by motley and striped
cornsnakes. Unlike those two pattern morphs, there is no tendency for
dark striping along the edges of the ventral scales, indicting a change in
distribution of ventral pigments. Rather, all pigmentation is simply
missing.
Banded
The genetic origins of this trait are unclear, and it may in
fact be a simple variant of the motley trait. However, it is distinct
enough in appearance to be discussed here. The dorsal blotches are
simply expanded down onto the lateral regions to give the appearance of a
banded snake, rather than blotched. The best specimens also appear to
lack the intermediate lateral markings. As this trait is fairly new to
the scene, it remains to be seen where it will lead.
Frosted
Appears to be a pattern trait in which pigmentation is concentrated
at the center of each scale, producing a pale speckled look. Has been
crossed to several color mutations. This trait can be seen to a degree in
many cornsnakes, but specimens which have been selected for this tendency are
very unique in appearance. Here's another relatively new genetic trait for
breeders to explore.
Wide-Stripe
A very new cornsnake
pattern variation, not yet fully understood. It appears as a very wide
mid-dorsal stripe, with slight wavy edges. Side patterns tend to be reduced
to thin vertical bars on many specimens. Mode of inheritance is uncertain at
this point, but our limited breeding results indicate this is may be a
dominant mutation. Thus far, all of our breedings between two wide-stripes
have yielded 100% wide-stripes. Breedings between wide-stripes and unrelated
anery and snow specimens have also yielded 100% wide-stripes. Currently very
few specimens exist.
Piebald
Generally termed 'Calico' in the trade, although there may in fact be two
or more mutations present in captive collections which fall under this
label. The first discovered incidence of piebaldism involves animals
which develop varying amounts of white scattered patches during later years of
life. These patches are associated with some form of cellular disruption
appearing as blistered deformities of the scales. Breeders are currently
attempting to see if this negative trait can be separated from the 'desirable'
one.
A few recent hatchlings have appeared to be
true piebald specimens, in which large areas are devoid of pigment at
birth. Future breeding projects will determine if this trait is genetically
inheritable or simply a developmental anomaly. The parallel trait in the
Ball Python (Python regius) was proven inheritable in the summer of
1999...
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